K. J. H. Wong, H.-T. Shih, and B. K. K. Chan, “The ghost crab Ocypode mortoni George, 1982 (Crustacea: Decapoda: Ocypodidae): redescription, distribution at its type locality, and the phylogeny of East Asian Ocypode species”, Zootaxa, vol. 3550, no. 1, pp. 71–87, Nov. 2012.