T. Lackner and Y. Gomy, “Review of the genus Saprinus Erichson, 1834 from Madagascar and adjacent islands with description of a new species (Coleoptera: Histeridae: Saprininae) Third contribution to the knowledge of the Histeridae of Madagascar”, Zootaxa, vol. 4083, no. 2, pp. 151–200, Feb. 2016.