A. R. Pérez-Asso, R. Núñez-Aguila, and J. A. Genaro, “Morphology and COI barcodes reveal four new species in the lycieus group of Calisto (Lepidoptera, Nymphalidae, Satyrinae)”, Zootaxa, vol. 4170, no. 3, pp. 401–450, Sep. 2016.