J. M. Wingert, J. Ferrer, and L. R. Malabarba, “Review of the Odontesthes perugiae species group from Río de La Plata drainage, with the description of a new species (Atherinomorpha: Atherinopsidae)”, Zootaxa, vol. 4250, no. 6, pp. 501–528, Apr. 2017.