L. L. Zupolini, T. Magalhães, L. G. Pileggi, and F. L. Mantelatto, “Taxonomic revision of the speckled crabs, genus Arenaeus Dana, 1851 (Brachyura: Portunidae) based on morphological and molecular data”, Zootaxa, vol. 4273, no. 3, pp. 362–380, Jun. 2017.