M. D. Escalona S., F. A. Juncá, A. A. Giaretta, A. J. Crawford, and E. L. Marca, “Contrasting genetic, acoustic, and morphological differentiation in two closely related gladiator frogs (Hylidae: Boana) across a common Neotropical landscape”, Zootaxa, vol. 4609, no. 3, pp. 519–547, May 2019.