D. R. Maddison, L. Toledano, S. Sallenave, and S. Roig-Juñent, “Phylogenetic relationships of the South American ground beetle subgenus Chilioperyphus Jeannel (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidion Latreille)”, Zootaxa, vol. 3636, no. 4, pp. 547–560, Apr. 2013.