R. A. Johnson, R. P. Overson, and C. S. Moreau, “A New Species of Seed-harvester Ant, Pogonomyrmex hoelldobleri (Hymenoptera: Formicidae), from the Mohave and Sonoran Deserts of North America”, Zootaxa, vol. 3646, no. 3, pp. 201–227, May 2013.