A. L. Netto-Ferreira and M. M. F. Marinho, “New species of Pyrrhulina (Ostariophysi: Characiformes: Lebiasinidae) from the Brazilian Shield, with comments on a putative monophyletic group of species in the genus”, Zootaxa, vol. 3664, no. 3, pp. 369–376, May 2013.