A. Rakotoarison, A. Crottini, J. Müller, M.-O. Rödel, F. Glaw, and M. Vences, “Revision and phylogeny of narrow-mouthed treefrogs (Cophyla) from northern Madagascar: integration of molecular, osteological, and bioacoustic data reveals three new species”, Zootaxa, vol. 3937, no. 1, pp. 61–89, Mar. 2015.