T. C. Macrae, “Agrilus (s. str.) betulanigrae MacRae (Coleoptera: Buprestidae: Agrilini), a new species from North America, with comments on subgeneric placement and a key to the otiosus species-group in North America”, Zootaxa, vol. 380, no. 1, pp. 1–9, Dec. 2003.