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P. J. Bartels, G. Pilato, O. Lisi, and D. R. Nelson, “Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee/North Carolina, USA (North America): two new species and six new records”, Zootaxa, vol. 2022, no. 1, pp. 45–57, Feb. 2009.