M.-O. Rödel, A. Schmitz, O. S. Pauwels, and W. Böhme, “Revision of the genus Werneria Poche, 1903, including the descriptions of two new species from Cameroon and Gabon (Amphibia: Anura: Bufonidae)”, Zootaxa, vol. 720, no. 1, pp. 1–28, Nov. 2004.