P. J. C. Russell, J. E. Pateman, A. V. Gagarina, and V. A. Lukhtanov, “Investigations into the Melitaea ornata species complex in the Levant: M. telona and the newly erected species Melitaea klili Benyamini, 2021 (Lepidoptera: Nymphalidae)”, Zootaxa, vol. 5285, no. 1, pp. 187–195, May 2023.