A. M. Shapovalov and M. F. Bagaturov, “A review of the subgenus Mesolethrus Nikolajev, 2003 of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae: Lethrinae) with description of a new species”, Zootaxa, vol. 5351, no. 4, pp. 435–452, Sep. 2023.