F. E. Mofrad, J. Muilwijk, A. A. Talebi, and S. Serri, “The ground beetle genus Zuphium and subgenus Chlaenius (Trichochlaenius) (Coleoptera: Carabidae) from Iran, with the description of two new species”, Zootaxa, vol. 5729, no. 4, pp. 566–580, Dec. 2025.