M. D. Barria, J. Clavijo-Bustos, and M. Seidel, “A synopsis and new species of Bolax Fischer von Waldheim, 1829 (Coleoptera: Scarabaeidae: Rutelinae: Geniatini) from Central America”, Zootaxa, vol. 5752, no. 2, pp. 276–284, Jan. 2026.