L. Beati, Émilie Bouhsira, C. Berlan, E. Liénard, A. A. Guglielmone, and S. Nava, “Type specimens of ixodid ticks (Acari: Ixodidae) in the ‘Neumann Collection’ of the National Veterinary School of Toulouse, France”, Zootaxa, vol. 5760, no. 2, pp. 209–224, Feb. 2026.