P. Urban, “A new species of Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Onthophagini) in the subgenus Gibbonthophagus Balthasar, 1935 from the mountains of north-west Yunnan, China”, Zootaxa, vol. 5837, no. 3, pp. 498–514, Jun. 2026.