L. A. Coloma, W. E. Duellman, A. A. C., S. R. Ron, A. Terán-Valdez, and J. M. Guayasamin, “Five new (extinct?) species of Atelopus (Anura: Bufonidae) from Andean Colombia, Ecuador, and Peru”, Zootaxa, vol. 2574, no. 1, pp. 1–54, Aug. 2010.