Brown, J. L., E. Twomey, A. Amézquita, M. B. de Souza, J. P. Caldwell, S. Lötters, R. Von May, P. R. Melo-Sampaio, D. Mejía-Vargas, P. Perez-Peña, M. Pepper, E. H. Poelman, M. Sanchez-Rodriguez, and K. Summers. “A Taxonomic Revision of the Neotropical Poison Frog Genus Ranitomeya (Amphibia: Dendrobatidae)”. Zootaxa, vol. 3083, no. 1, Oct. 2011, pp. 1–120, doi:10.11646/zootaxa.3083.1.1.