Gareca, Y., and P. Blandin. “Morpho (Morpho) helenor (Cramer) (Lepidoptera, Nymphalidae, Morphinae) in Bolivia: Geographical Distribution and Ecological Plasticity, With a Description of a New Subspecies”. Zootaxa, vol. 3130, no. 1, Dec. 2011, pp. 30–56, doi:10.11646/zootaxa.3130.1.2.