Hutter, C. R., and J. M. Guayasamin. “A New Cryptic Species of Glassfrog (Centrolenidae: Nymphargus) from Reserva Las Gralarias, Ecuador”. Zootaxa, vol. 3257, no. 1, Apr. 2012, pp. 1–21, doi:10.11646/zootaxa.3257.1.1.