Jansen, M., and A. Schulze. “Molecular, Morphology and Bioacoustic Data Suggest Bolivian Distribution of a Large Species of the Leptodactylus Pentadactylus Group (Amphibia: Anura: Leptodactylidae)”. Zootaxa, vol. 3307, no. 1, May 2012, pp. 35–47, doi:10.11646/zootaxa.3307.1.2.