Chen, W.-J., and R. L. Mayden. “Phylogeny of Suckers (Teleostei: Cypriniformes: Catostomidae): Further Evidence of Relationships Provided by the Single-Copy Nuclear Gene IRBP2”. Zootaxa, vol. 3586, no. 1, Dec. 2012, pp. 195–210, doi:10.11646/zootaxa.3586.1.18.