Nunes, R. V., and F. Z. Vaz-De-Mello. “A New Species of Holocephalus Hope from Paraguay (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini S. Str.)”. Zootaxa, vol. 4136, no. 2, July 2016, pp. 397–400, doi:10.11646/zootaxa.4136.2.12.