Ponssa, M. L. “On the Osteology of a Distinctive Specie of the Genus Leptodactylus: Leptodactylus Laticeps (Boulenger, 1917) (Anura, Leptodactylidae)”. Zootaxa, vol. 1188, no. 1, Apr. 2006, pp. 23–36, doi:10.11646/zootaxa.1188.1.2.