Naka, T., and M. Maruyama. “Aphaenogaster gamagumayaa Sp. nov.: The First Troglobiotic Ant from Japan (Hymenoptera: Formicidae: Myrmicinae)”. Zootaxa, vol. 4450, no. 1, July 2018, pp. 135–141, doi:10.11646/zootaxa.4450.1.10.