de Souza, L. S., D. C. Taphorn, and J. W. Armbruster. “Review of Ancistrus (Siluriformes: Loricariidae) from the Northwestern Guiana Shield, Orinoco Andes, and Adjacent Basins With Description of Six New Species”. Zootaxa, vol. 4552, no. 1, Feb. 2019, pp. 1–67, doi:10.11646/zootaxa.4552.1.1.