Tuzovskij, P. V. “New Water Mite Species of the Genus Piona Koch, 1842 (Acari, Hydrachnidia: Pionidae) from Northeastern Russia”. Zootaxa, vol. 3664, no. 3, May 2013, pp. 321–334, doi:10.11646/zootaxa.3664.3.3.