Angulo, A., and J. M. Gracian-Negrete. “A New Species of Brycon (Characiformes: Characidae) from Nicaragua and Costa Rica, With a Key to the Lower Mesoamerican Species of the Genus”. Zootaxa, vol. 3731, no. 2, Oct. 2013, pp. 255–266, doi:10.11646/zootaxa.3731.2.6.