Erwin, T. L. “The Beetle Family Carabidae of Costa Rica: Twenty-Nine New Species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)”. Zootaxa, vol. 119, no. 1, Dec. 2002, pp. 1–68, doi:10.11646/zootaxa.119.1.1.