Köhler, J., F. Glaw, M. Pabijan, and M. Vences. “Integrative Taxonomic Revision of Mantellid Frogs of the Genus Aglyptodactylus (Anura: Mantellidae)”. Zootaxa, vol. 4006, no. 3, Aug. 2015, pp. 401–438, doi:10.11646/zootaxa.4006.3.1.