Ahyong, S. T., and P. K. Ng. “New Species of Pinnotherid Crabs from Southeast Asia and Papua New Guinea (Crustacea: Decapoda: Brachyura)”. Zootaxa, vol. 4816, no. 3, July 2020, pp. 333–349, doi:10.11646/zootaxa.4816.3.4.