Brown, J. L., E. Twomey, M. Pepper, and M. S. Rodriguez. “Revision of the Ranitomeya fantastica Species Complex With Description of Two New Species from Central Peru (Anura: Dendrobatidae)”. Zootaxa, vol. 1823, no. 1, July 2008, pp. 1–24, doi:10.11646/zootaxa.1823.1.1.