Bartels, P. J., G. Pilato, O. Lisi, and D. R. Nelson. “Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee/North Carolina, USA (North America): Two New Species and Six New Records”. Zootaxa, vol. 2022, no. 1, Feb. 2009, pp. 45–57, doi:10.11646/zootaxa.2022.1.4.