Rödel, M.-O., A. Schmitz, O. S. Pauwels, and W. Böhme. “Revision of the Genus Werneria Poche, 1903, Including the Descriptions of Two New Species from Cameroon and Gabon (Amphibia: Anura: Bufonidae)”. Zootaxa, vol. 720, no. 1, Nov. 2004, pp. 1-28, doi:10.11646/zootaxa.720.1.1.