Harrison, J. D. G. “A Taxonomic Revision of the African Leaf Chafer Genus Asthenopholis Brenske, 1898 (Coleoptera: Scarabaeidae: Melolonthinae): A SEM Study”. Zootaxa, vol. 2225, no. 1, Sept. 2009, pp. 1–48, doi:10.11646/zootaxa.2225.1.1.