Deschodt, C. M., and C. L. Sole. “A New Genus and Species in the Diverse Dung Beetle Tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa”. Zootaxa, vol. 5375, no. 2, Nov. 2023, pp. 279-84, doi:10.11646/zootaxa.5375.2.9.