Deschodt, C. M., and C. L. Sole. “A New and Unusually Ornate Southern African Dung Beetle Species in d’Orbigny’s Onthophagus Group 18 (Coleoptera: Scarabaeidae: Onthophagini)”. Zootaxa, vol. 5493, no. 2, Aug. 2024, pp. 186-94, doi:10.11646/zootaxa.5493.2.7.