Samyn, Y., and C. Massin. “A Tale of Three Oceans—Taxonomy of the Holothuria (Thymiosycia) arenicola Semper, 1868 Complex (Echinodermata: Holoturoidea: Holothuriidae)”. Zootaxa, vol. 5507, no. 4, Sept. 2024, pp. 501-33, doi:10.11646/zootaxa.5507.4.1.