Selis, M., and R. W. Longair. “New Data on the Genus Belonogaster De Saussure, With Establishment of the atrata Species-Group (Hymenoptera: Vespidae: Polistinae)”. Zootaxa, vol. 5692, no. 2, Sept. 2025, pp. 353-68, doi:10.11646/zootaxa.5692.2.8.