Génier, F. “Three New Species of Onthophagus Latreille, 1802 from Bioko Island, Equatorial Guinea, Belonging to d’Orbigny’s Group IX, With a Taxonomic Note on O. denudatus D’Orbigny, 1902 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)”. Zootaxa, vol. 5831, no. 3, June 2026, pp. 413-26, doi:10.11646/zootaxa.5831.3.6.