Hutter, Carl R., and Juan M. Guayasamin. “A New Cryptic Species of Glassfrog (Centrolenidae: Nymphargus) from Reserva Las Gralarias, Ecuador”. Zootaxa 3257, no. 1 (April 4, 2012): 1–21. Accessed April 9, 2025. https://mapress.com/zt/article/view/zootaxa.3257.1.1.