Arce-Pérez, Roberto, and Miguel Angel Morón. “New Species and New Records of Species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua”. Zootaxa 1012, no. 1 (June 28, 2005): 23–37. Accessed April 24, 2025. https://mapress.com/zt/article/view/zootaxa.1012.1.3.