Yan, Zhuo, Ming Bai, and Dong Ren. “A New Fossil Hybosoridae (Coleoptera: Scarabaeoidea) from the Yixian Formation of China”. Zootaxa 3478, no. 1 (September 11, 2012): 201–204. Accessed February 9, 2026. https://mapress.com/zt/article/view/zootaxa.3478.1.21.