López-García, Margarita M., Héctor J. Gasca-Álvarez, Ronald D. Cave, and Germán Amat-García. “An Annotated Checklist of the New World Pentodontine Scarab Beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) ”. Zootaxa 4170, no. 3 (September 26, 2016): 491–509. Accessed April 22, 2025. https://mapress.com/zt/article/view/zootaxa.4170.3.4.