Iwan, Dariusz. “Species Groups in the Genus Atrocrates Koch, 1956 (Coleoptera: Tenebrionidae: Pedinini)”. Zootaxa 4175, no. 6 (October 18, 2016): 571–587. Accessed April 22, 2025. https://mapress.com/zt/article/view/zootaxa.4175.6.5.