Rakotonirina, Jean Claude, Sándor Csősz, and Brian L. Fisher. “Taxonomic Revision of the Malagasy Camponotus grandidieri and niveosetosus Species Groups (Hymenoptera, Formicidae) Using Qualitative and Quantitative Morphology”. Zootaxa 4238, no. 2 (March 2, 2017): 203–245. Accessed April 3, 2025. https://mapress.com/zt/article/view/zootaxa.4238.2.2.