Valencia, Jorge H., Francisco Valladares-Suntasig, Luis Tipantiza-Tuguminago, and Manuel R. Dueñas. “A New Species of Terrestrial-Breeding Frog of the Genus Pristimantis (Anura: Terrarana: Craugastoridae) from the Eastern Andean Slopes of the Southern Ecuador”. Zootaxa 4658, no. 3 (August 23, 2019): 509–525. Accessed April 12, 2025. https://mapress.com/zt/article/view/zootaxa.4658.3.4.